CAMK1A

All methods over the ±5 window, side by side.

PSPA

CDDM

MLP-attr

Normalized values from positional scanning peptide array

PSPA measures pT, so at flanking positions the pS row is a duplicate of the pT row. At position 0 the acceptor identity (pS, pT, pY) is genuine.

Position-wise Probabilities

Log-Odds: Probabilities / STY Background

Position-wise Probabilities (±5 window)

Log-Odds (±5 window)

Motifs stratified by phospho-acceptor (S / T / Y)

K-means sub-motif clusters

Signed attribution PSSM distilled from the CDDM discriminative MLP (in-silico saturation mutagenesis). Letters above 0 are enriched, below are depleted; this is not a probability distribution.

MLP-attribution PSSM (±5, signed)

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substrate_uniprot	site	source	substrate_genes	site_seq
A0A2R8Y4L2	S95	Sugiyama	HNRNPA1L3 HNRNPA1P48	RPHKVDGRVVEPKRAVSREDsQRPDAHLTVKKIFVGGIKED
O00429	S637	SIGNOR	DNM1L DLP1 DRP1	PQKGHAVNLLDVPVPVARKLsAREQRDCEVIERLIKSYFLI
O14639	S452	Sugiyama	ABLIM1 ABLIM KIAA0059 LIMAB1	PtPsAEGyQDVRDRMIHRstsQGsINsPVySRHsYtPTTSR
O14974	S910	Sugiyama	PPP1R12A MBS MYPT1	yETssTsAGDRyDsLLGRsGsysYLEERKPYSSRLEKDDST
O43432	S1156	SIGNOR\|iPTMNet\|PSP	EIF4G3	KNDKPLPSATARPNTFMRGGssKDLLDNQsQEEQRREMLET
O43823	S323	Sugiyama	AKAP8 AKAP95	RKRKQFQLyEEPDtKLARVDsEGDFsENDDAAGDFRsGDEE
O60716	S252	Sugiyama	CTNND1 KIAA0384	sRVtRIEERyRPsMEGyRAPsRQDVyGPQPQVRVGGssVDL
O60841	S186	Sugiyama	EIF5B IF2 KIAA0741	EDEDNsKKIKERsRINssGEsGDEsDEFLQsRKGQKKNQKN
O95714	S2928	Sugiyama	HERC2	ILLGRIRAEEEDLAAVPFLAsDNEEEEDEKGNSGsLIRKKA
P00558	Y161	Sugiyama	PGK1 PGKA MIG10 OK/SW-cl.110	KAEPAKIEAFRAsLSKLGDVyVNDAFGtAHRAHssMVGVNL
P04075	S36	Sugiyama	ALDOA ALDA	LsDIAHRIVAPGKGILAADEstGsIAKRLQsIGtENtEENR
P04406	S241	Sugiyama	GAPDH GAPD CDABP0047 OK/SW-cl.12	IPELNGKLtGMAFRVPtANVsVVDLtCRLEKPAKyDDIKKV
P04792	S15	Sugiyama	HSPB1 HSP27 HSP28	______MTERRVPFsLLRGPsWDPFRDWyPHsRLFDQAFGL
P04792	S82	Sugiyama	HSPB1 HSP27 HSP28	AIEsPAVAAPAYsRALsRQLssGVsEIRHtADRWRVsLDVN
P05783	S53	SIGNOR\|ELM	KRT18 CYK18 PIG46	AsVyAGAGGsGsRIsVsRstsFRGGMGsGGLAtGIAGGLAG
P06733	Y189	Sugiyama	ENO1 ENO1L1 MBPB1 MPB1	MILPVGAANFREAMRIGAEVyHNLKNVIKEKyGKDAtNVGD
P06733	Y44	Sugiyama	ENO1 ENO1L1 MBPB1 MPB1	LFtsKGLFRAAVPsGAstGIyEALELRDNDKtRYMGKGVSK
P07205	Y161	Sugiyama	PGK2 PGKB	KAEPDKIEAFRAsLSKLGDVyVNDAFGtAHRAHSSMVGVNL
P07900	Y160	Sugiyama	HSP90AA1 HSP90A HSPC1 HSPCA	SAYLVAEKVTVITKHNDDEQyAWEssAGGsFtVRTDTGEPM
P08238	Y155	Sugiyama	HSP90AB1 HSP90B HSPC2 HSPC3 HSPCB	SAYLVAEKVVVITKHNDDEQyAWEssAGGsFtVRADHGEPI
P08238	Y301	Sugiyama	HSP90AB1 HSP90B HSPC2 HSPC3 HSPCB	ELNKtKPIWtRNPDDItQEEyGEFyKsLtNDWEDHLAVKHF
P08238	Y305	Sugiyama	HSP90AB1 HSP90B HSPC2 HSPC3 HSPCB	tKPIWtRNPDDItQEEyGEFyKsLtNDWEDHLAVKHFSVEG
P08708	S89	Sugiyama	RPS17 RPS17L	ISIKLQEEERERRDNyVPEVsALDQEIIEVDPDtKEMLKLL
P08708	Y84	Sugiyama	RPS17 RPS17L	GPVRGISIKLQEEERERRDNyVPEVsALDQEIIEVDPDtKE
P09104	Y44	Sugiyama	ENO2	LyTAKGLFRAAVPsGAstGIyEALELRDGDKQRYLGKGVLK
P09651	S95	Sugiyama	HNRNPA1 HNRPA1	RPHKVDGRVVEPKRAVsREDsQRPGAHLtVKKIFVGGIKED
P13929	Y44	Sugiyama	ENO3	LHtAKGRFRAAVPsGAstGIyEALELRDGDKGRYLGKGVLK
P17544	S44	Sugiyama	ATF7 ATFA	LAVHKHKHEMTLKFGPARtDsVIIADQtPtPtRFLKNCEEV
P17600	S9	SIGNOR\|iPTMNet	SYN1	____________MNYLRRRLsDSNFMANLPNGYMTDLQRPQ
P18206	S1002	Sugiyama	VCL	SKGNDIIAAAKRMALLMAEMsRLVRGGSGTKRALIQCAKDI
P18846	S63	SIGNOR\|iPTMNet	ATF1	DsSDsIGssQKAHGILARRPsYRKILKDLsSEDTRGRKGDG
P19474	S3	Sugiyama	TRIM21 RNF81 RO52 SSA1	__________________MAsAARLTMMWEEVTCPICLDPF
P22626	S189	Sugiyama	HNRNPA2B1 HNRPA2B1	IVLQKyHtINGHNAEVRKALsRQEMQEVQssRsGRGGNFGF
P23588	Y609	Sugiyama	EIF4B	sASKyAALsVDGEDENEGEDyAE__________________
P24534	S141	Sugiyama	EEF1B2 EEF1B EF1B	ERLAQYESKKAKKPALVAKssILLDVKPWDDEtDMAKLEEC
P24844	S20	PSP	MYL9 MLC2 MRLC1 MYRL2	_MssKRAKAKtTKKRPQRAtsNVFAMFDQSQIQEFKEAFNM
P26373	S77	Sugiyama	RPL13 BBC1 OK/SW-cl.46	PIVRCPTVRYHTKVRAGRGFsLEELRVAGIHKKVARTIGIs
P29475	S746	iPTMNet	NOS1	GTPTKRRAIGFKKLAEAVKFsAKLMGQAMAKRVKATILYAT
P29475	S852	SIGNOR\|iPTMNet	NOS1	EMRHPNSVQEERKSYKVRFNsVSSYSDsQKSSGDGPDLRDN
P32119	T142	Sugiyama	PRDX2 NKEFB TDPX1	EGIAYRGLFIIDGKGVLRQItVNDLPVGRsVDEALRLVQAF
P35968	Y951	Sugiyama	KDR FLK1 VEGFR2	KRNEFVPYKTKGARFRQGKDyVGAIPVDLKRRLDSITSSQS
P38646	T398	Sugiyama	HSPA9 GRP75 HSPA9B mt-HSP70	SDIGEVILVGGMTRMPKVQQtVQDLFGRAPsKAVNPDEAVA
P39019	S59	PSP	RPS19	LAKHKELAPyDENWFytRAAstARHLYLRGGAGVGsMTKIY
P39019	Y48	Sugiyama	RPS19	LKVPEWVDtVKLAKHKELAPyDENWFytRAAstARHLYLRG
P39019	Y54	Sugiyama	RPS19	VDtVKLAKHKELAPyDENWFytRAAstARHLYLRGGAGVGs
P40818	S719	Sugiyama	USP8 KIAA0055 UBPY	KPQIPAERDREPSKLKRsyssPDItQAIQEEEKRKPTVtPT
P43243	S188	Sugiyama	MATR3 KIAA0723	EPPyRVPRDDWEEKRHFRRDsFDDRGPsLNPVLDyDHGsRs
P43487	S60	Sugiyama	RANBP1	TLEEDEEELFKMRAKLFRFAsENDLPEWKERGTGDVKLLKH
P46013	T1764	Sugiyama	MKI67	LVDtPtssKPQPKRSLRKADtEEEFLAFRKQTPSAGKAMHt
P46527	T157	SIGNOR\|PSP	CDKN1B KIP1 p27	PsDsQTGLAEQCAGIRKRPAtDDSSTQNKRANRTEENVsDG
P46527	T198	SIGNOR\|PSP	CDKN1B KIP1 p27	sPNAGSVEQtPKKPGLRRRQt____________________
P46777	S224	Sugiyama	RPL5 MSTP030	VADyMRyLMEEDEDAyKKQFsQyIKNsVtPDMMEEMyKKAH
P49327	T2215	Sugiyama	FASN FAS	DEAsELACPtPKEDGLAQQQtQLNLRSLLVNPEGPtLMRLN
P49757	S276	SIGNOR	NUMB C14orf41	NNPHAIPRRHAPIEQLARQGsFRGFPALsQKMSPFKRQLsL
P52943	S115	Sugiyama	CRIP2 CRP2	AARAEERKAsGPPKGPsRAssVttFtGEPNtCPRCSKKVYF
P53999	S118	Sugiyama	SUB1 PC4 RPO2TC1	PGRKGIsLNPEQWsQLKEQIsDIDDAVRKL___________
P60709	T303	Sugiyama	ACTB	MKCDVDIRKDLyANtVLsGGttMyPGIADRMQKEItALAPs
P60709	Y306	Sugiyama	ACTB	DVDIRKDLyANtVLsGGttMyPGIADRMQKEItALAPstMK
P61604	Y88	Sugiyama	HSPE1	GDKVLLPEyGGtKVVLDDKDyFLFRDGDILGKyVD______
P62273	Y7	Sugiyama	RPS29	______________MGHQQLyWsHPRKFGQGSRSCRVCSNR
P63220	S65	Sugiyama	RPS21	RFNGQFKtyAICGAIRRMGEsDDsILRLAKADGIVSKNF__
P63220	Y53	Sugiyama	RPS21	QMNVAEVDKVTGRFNGQFKtyAICGAIRRMGEsDDsILRLA
P63261	T303	Sugiyama	ACTG1 ACTG	MKCDVDIRKDLyANtVLsGGttMyPGIADRMQKEItALAPs
P63261	Y306	Sugiyama	ACTG1 ACTG	DVDIRKDLyANtVLsGGttMyPGIADRMQKEItALAPstMK
P67809	S102	Sugiyama	YBX1 NSEP1 YB1	EDVFVHQtAIKKNNPRKyLRsVGDGEtVEFDVVEGEKGAEA
P67809	T108	Sugiyama	YBX1 NSEP1 YB1	QtAIKKNNPRKyLRsVGDGEtVEFDVVEGEKGAEAANVtGP
P68363	Y432	Sugiyama	TUBA1B	GMEEGEFSEAREDMAALEKDyEEVGVDsVEGEGEEEGEEy_
P83731	T83	Sugiyama	RPL24	QsEEIQKKRTRRAVKFQRAItGAsLADIMAKRNQKPEVRKA
Q00587	S192	Sugiyama	CDC42EP1 BORG5 CEP1 MSE55	PHDRDRDGsFPsEPGLRRsDsLLsFRLDLDLGPSLLSELLG
Q01082	S1388	Sugiyama	SPTBN1 SPTB2	TTQTKAQRLFDANKAELFtQsCADLDKWLHGLESQIQSDDy
Q01105	S133	Sugiyama	SET	DEEALHYLTRVEVtEFEDIKsGyRIDFyFDENPyFENKVLS
Q01105	Y140	Sugiyama	SET	LTRVEVtEFEDIKsGyRIDFyFDENPyFENKVLSKEFHLNE
Q01105	Y146	Sugiyama	SET	tEFEDIKsGyRIDFyFDENPyFENKVLSKEFHLNEsGDPss
Q05209	S435	Sugiyama	PTPN12	LPGKNESTIEQIDKKLERNLsFEIKKVPLQEGPKsFDGNtL
Q05469	S855	PSP	LIPE	SGRKSQKMSEPIAEPMRRsVsEAALAQPQGPLGTDSLKNLT
Q06830	T143	Sugiyama	PRDX1 PAGA PAGB TDPX2	EGIsFRGLFIIDDKGILRQItVNDLPVGRsVDEtLRLVQAF
Q09472	S89	ELM	EP300 P300	GMVQDAASKHKQLSELLRsGssPNLNMGVGGPGQVMAsQAQ
Q13085	S25	ELM	ACACA ACAC ACC1 ACCA	sPLAQPLELNQHSRFIIGsVsEDNsEDEISNLVKLDLLEEK
Q14012	S324	Sugiyama	CAMK1	KQAFNAtAVVRHMRKLQLGtsQEGQGQtAsHGELLtPVAGG
Q14012	S333	Sugiyama	CAMK1	VRHMRKLQLGtsQEGQGQtAsHGELLtPVAGGPAAGCCCRD
Q14012	S363	Sugiyama	CAMK1	GGPAAGCCCRDCCVEPGtELsPtLPHQL_____________
Q14012	T177	SIGNOR\|ELM\|iPTMNet	CAMK1	KIMISDFGLSKMEDPGSVLstACGTPGYVAPEVLAQKPYSK
Q14012	T310	Sugiyama	CAMK1	VSEQIKKNFAKSKWKQAFNAtAVVRHMRKLQLGtsQEGQGQ
Q14012	T323	Sugiyama	CAMK1	WKQAFNAtAVVRHMRKLQLGtsQEGQGQtAsHGELLtPVAG
Q14012	T331	Sugiyama	CAMK1	AVVRHMRKLQLGtsQEGQGQtAsHGELLtPVAGGPAAGCCC
Q14012	T339	Sugiyama	CAMK1	LQLGtsQEGQGQtAsHGELLtPVAGGPAAGCCCRDCCVEPG
Q14012	T365	Sugiyama	CAMK1	PAAGCCCRDCCVEPGtELsPtLPHQL_______________
Q14568	Y160	Sugiyama	HSP90AA2P HSP90AA2 HSPC2 HSPCAL3	SAYLVAEKVTVITKHNDDEQyAWEssAGGsFtVRTDTGERM
Q16566	T200	ELM	CAMK4 CAMK CAMK-GR CAMKIV	PLKIADFGLsKIVEHQVLMKtVCGTPGYCAPEILRGCAYGP
Q32P51	S95	Sugiyama	HNRNPA1L2 HNRNPA1L	TPHKVDGRVVEPKRAVsREDsQRPGAHLtVKKIFVGGIKED
Q5UIP0	S1688	Sugiyama	RIF1	PVPESNLRTRNAIKRLHKRDsFDNCsLGESSKIGIsDIssL
Q71U36	Y432	Sugiyama	TUBA1A TUBA3	GMEEGEFSEAREDMAALEKDyEEVGVDsVEGEGEEEGEEy_
Q7Z417	S652	Sugiyama	NUFIP2 KIAA1321 PIG1	NtLLGsAKEQRYQRGLERNDsWGsFDLRAAIVYHTKEMEsI
Q86V81	Y250	Sugiyama	ALYREF ALY BEF THOC4	AGRNSKQQLsAEELDAQLDAyNARMDts_____________
Q8IU85	T313	Sugiyama	CAMK1D CAMKID	VSAQIRKNFAKSKWRQAFNAtAVVRHMRKLHLGssLDssNA
Q8NFH8	S631	Sugiyama	REPS2 POB1	AIQTAIRKNKEANAVLARLNsELQQQLKEVHQERIALENQL
Q8WVC0	T477	Sugiyama	LEO1 RDL	FDVYKAPLQGDHNHLFIRQGtGLQGQAVFKTKLTFRPHstD
Q96D71	S767	Sugiyama	REPS1	AIQAsIRRNKETNTVLARLNsELQQQLKDVLEERISLEVQL
Q99618	S94	Sugiyama	CDCA3 C8 GRCC8 TOME1	ARtPMKTSsGDPPsPLVKQLsEVFETEDSKSNLPPEPVLPP
Q9BQE3	Y432	Sugiyama	TUBA1C TUBA6	GMEEGEFSEAREDMAALEKDyEEVGADsADGEDEGEEy___
Q9BRS2	T509	Sugiyama	RIOK1 RIO1	QVEERtCsDsEDIGssECsDtDSEEQGDHARPKKHTTDPDI
Q9NP62	S177	PSP	GCM1 GCMA	PETKLEAEARRAMKKVNtAPssVSLSLKGSTETRSLPGETQ
Q9NP62	S178	PSP	GCM1 GCMA	ETKLEAEARRAMKKVNtAPssVSLSLKGSTETRSLPGETQS
Q9NP62	S269	PSP	GCM1 GCMA	DLTDQTSTVDPMKLYEKRKLsSSRTYsSGDLLPPSASGVYS
Q9NP62	S47	SIGNOR\|PSP	GCM1 GCMA	VKKTDWFQEWPDSYAKHIYSsEDKNAQRHLSSWAMRNTNNH
Q9NP62	T174	PSP	GCM1 GCMA	HPKPETKLEAEARRAMKKVNtAPssVSLSLKGSTETRSLPG
Q9NY27	S311	Sugiyama	PPP4R2 SBBI57	CTRQHCtEEDEEEDEEEEEEsFMtsREMIPERKNQEKESDD
Q9NY27	S315	Sugiyama	PPP4R2 SBBI57	HCtEEDEEEDEEEEEEsFMtsREMIPERKNQEKESDDALTV
Q9NY27	T297	Sugiyama	PPP4R2 SBBI57	EAssssQDKDKDSRCTRQHCtEEDEEEDEEEEEEsFMtsRE
Q9NY27	T314	Sugiyama	PPP4R2 SBBI57	QHCtEEDEEEDEEEEEEsFMtsREMIPERKNQEKESDDALT
Q9UKV0	S220	PSP	HDAC9 HDAC7 HDAC7B HDRP KIAA0744 MITR	KYTLPGAQDAKDDFPLRKTAsEPNLKVRSRLKQKVAERRss
Q9UKV0	S451	PSP	HDAC9 HDAC7 HDAC7B HDRP KIAA0744 MITR	SPGIRGTHKLPRHRPLNRtQsAPLPQSTLAQLVIQQQHQQF
Q9UQL6	S259	SIGNOR\|iPTMNet\|PSP	HDAC5 KIAA0600	KLPLPGPYDSRDDFPLRKTAsEPNLKVRSRLKQKVAERRss
Q9UQL6	S498	SIGNOR\|iPTMNet\|PSP	HDAC5 KIAA0600	ATSMRTVGKLPRHRPLSRtQssPLPQSPQALQQLVMQQQHQ
Q9Y570	S15	SIGNOR\|PSP	PPME1 PME1 PP2593 PRO0750	______MSALEKSMHLGRLPsRPPLPGsGGsQsGAKMRMGP