CLK3

All methods over the ±5 window, side by side.
PSPA
CDDM
MLP-attr
Normalized values from positional scanning peptide array
PSPA measures pT, so at flanking positions the pS row is a duplicate of the pT row. At position 0 the acceptor identity (pS, pT, pY) is genuine.
Position-wise Probabilities
Log-Odds: Probabilities / STY Background
Position-wise Probabilities (±5 window)
Log-Odds (±5 window)
Motifs stratified by phospho-acceptor (S / T / Y)
K-means sub-motif clusters
Signed attribution PSSM distilled from the CDDM discriminative MLP (in-silico saturation mutagenesis). Letters above 0 are enriched, below are depleted; this is not a probability distribution.
MLP-attribution PSSM (±5, signed)
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substrate_uniprot site source substrate_genes site_seq
A0A2R8Y4L2 S95 Sugiyama HNRNPA1L3 HNRNPA1P48 RPHKVDGRVVEPKRAVSREDsQRPDAHLTVKKIFVGGIKED
O00488 S93 Sugiyama ZNF593 ZT86 STNLKTHFRSKDHKKRLKQLsVEPysQEEAERAAGMGSYVP
O43823 S124 Sugiyama AKAP8 AKAP95 SKEGGRGGsGGGGEGIQDREssFRFQPFESYDSRPCLPEHN
O43852 T65 Sugiyama CALU FDyDHDAFLGAEEAKtFDQLtPEEsKERLGKIVSKIDGDKD
O60292 S158 Sugiyama SIPA1L3 KIAA0545 SPAL3 FHRLSRRRsKDVEFQDGWPRsPGRAFLPLRHRsssEItLSE
O60343 S588 Sugiyama TBC1D4 AS160 KIAA0603 tssLENIFSRGANRMRGRLGsVDsFERsNsLASEKDYsPGD
O60841 S186 Sugiyama EIF5B IF2 KIAA0741 EDEDNsKKIKERsRINssGEsGDEsDEFLQsRKGQKKNQKN
P04075 S39 Sugiyama ALDOA ALDA IAHRIVAPGKGILAADEstGsIAKRLQsIGtENtEENRRFy
P04264 S21 Sugiyama KRT1 KRTA MSRQFSSRSGYRSGGGFSSGsAGIINYQRRTTSSSTRRSGG
P04792 S82 Sugiyama HSPB1 HSP27 HSP28 AIEsPAVAAPAYsRALsRQLssGVsEIRHtADRWRVsLDVN
P05386 S101 Sugiyama RPLP1 RRP1 APstAAAPAEEKKVEAKKEEsEEsDDDMGFGLFD_______
P05387 S102 Sugiyama RPLP2 D11S2243E RPP2 PAAGsAPAAAEEKKDEKKEEsEEsDDDMGFGLFD_______
P07237 S449 Sugiyama P4HB ERBA2L PDI PDIA1 PO4DB ANEVEAVKVHsFPtLKFFPAsADRtVIDyNGERtLDGFKKF
P08195 S607 Sugiyama SLC3A2 MDU1 ASLPAKADLLLstQPGREEGsPLELERLKLEPHEGLLLRFP
P09651 S95 Sugiyama HNRNPA1 HNRPA1 RPHKVDGRVVEPKRAVsREDsQRPGAHLtVKKIFVGGIKED
P13639 S584 Sugiyama EEF2 EF2 DHACIPIKKSDPVVsyREtVsEEsNVLCLsKsPNKHNRLyM
P13861 S99 Sugiyama PRKAR2A PKR2 PRKAR2 EsEEDEDLEVPVPsRFNRRVsVCAEtyNPDEEEEDTDPRVI
P14314 S126 Sugiyama PRKCSH G19P1 yNsGVICENtCKEKGRKEREsLQQMAEVtREGFRLKKILIE
P14625 S64 Sugiyama HSP90B1 GRP94 HSPC4 TRA1 tDDEVVQREEEAIQLDGLNAsQIRELREKSEKFAFQAEVNR
P15311 S539 Sugiyama EZR VIL2 TEAEKNERVQRQLLtLssELsQARDENKRTHNDIIHNENMR
P17480 S546 Sugiyama UBTF UBF UBF1 EKLMWIKKAAEDQKRYERELsEMRAPPAATNSSKKMKFQGE
P17812 S573 Sugiyama CTPS1 CTPS HYLQKGCRLsPRDtysDRsGsssPDsEItELKFPsINHD__
P18621 S5 Sugiyama RPL17 ________________MVRysLDPENPtKsCKSRGSNLRVH
P21333 S1084 Sugiyama FLNA FLN FLN1 PTKPSKVKAFGPGLQGGsAGsPARFtIDTKGAGTGGLGLTV
P22234 S27 Sugiyama PAICS ADE2 AIRC PAIS LNIGKKLYEGKtKEVyELLDsPGKVLLQsKDQItAGNAARK
P26038 S576 Sugiyama MSN YKtLRQIRQGNTKQRIDEFEsM___________________
P26373 S77 Sugiyama RPL13 BBC1 OK/SW-cl.46 PIVRCPTVRYHTKVRAGRGFsLEELRVAGIHKKVARTIGIs
P29692 S133 Sugiyama EEF1D EF1D NVLEKssPGHRAtAPQtQHVsPMRQVEPPAKKPAtPAEDDE
P30101 T485 Sugiyama PDIA3 ERP57 ERP60 GRP58 KKyEGGRELsDFIsyLQREAtNPPVIQEEKPKKKKKAQEDL
P35613 S279 Sugiyama BSG UNQ6505/PRO21383 TDSEDKALMNGSESRFFVsssQGRSELHIENLNMEADPGQy
P36578 S295 Sugiyama RPL4 RPL1 sNyNLPMHKMINtDLsRILKsPEIQRALRAPRKKIHRRVLK
P37802 S185 Sugiyama TAGLN2 KIAA0120 CDABP0035 NQLQEGKNVIGLQMGtNRGAsQAGMtGyGMPRQIL______
P39023 S13 Sugiyama RPL3 OK/SW-cl.32 ________MSHRKFSAPRHGsLGFLPRKRSSRHRGKVKSFP
P39023 S265 Sugiyama RPL3 OK/SW-cl.32 HRGLRKVACIGAWHPARVAFsVARAGQKGYHHRTEINKKIY
P41252 S1047 Sugiyama IARS1 IARS ESHTEFIFTTIKAPLKPYPVsPsDKVLIQEKTQLKGSELEI
P42166 S159 Sugiyama TMPO LAP2 RKLYEKKLLKLREQGtEsRsstPLPtIsssAENtRQNGsND
P42167 S159 Sugiyama TMPO LAP2 RKLYEKKLLKLREQGTESRsstPLPTISsSAENTRQNGSND
P43243 S188 Sugiyama MATR3 KIAA0723 EPPyRVPRDDWEEKRHFRRDsFDDRGPsLNPVLDyDHGsRs
P43243 S195 Sugiyama MATR3 KIAA0723 RDDWEEKRHFRRDsFDDRGPsLNPVLDyDHGsRsQEsGyyD
P53999 S56 Sugiyama SUB1 PC4 RPO2TC1 PVKKQKTGETsRALsssKQssssRDDNMFQIGKMRYVSVRD
P54105 S102 Sugiyama CLNS1A CLCI ICLN VNAKFEEEsKEPVADEEEEDsDDDVEPITEFRFVPSDKSAL
P55081 S132 Sugiyama MFAP1 VVGEsDsEVEGDAWRMEREDssEEEEEEIDDEEIERRRGMM
P62249 S9 Sugiyama RPS16 ____________MPsKGPLQsVQVFGRKKTATAVAHCKRGN
P62277 S12 Sugiyama RPS13 _________MGRMHAPGKGLsQsALPyRRsVPtWLKLTsDD
P62277 S14 Sugiyama RPS13 _______MGRMHAPGKGLsQsALPyRRsVPtWLKLTsDDVK
P62280 T15 Sugiyama RPS11 ______MADIQTERAyQKQPtIFQNKKRVLLGEtGKEKLPR
P62917 S14 Sugiyama RPL8 _______MGRVIRGQRKGAGsVFRAHVKHRKGAARLRAVDF
P62942 S9 Sugiyama FKBP1A FKBP1 FKBP12 ____________MGVQVEtIsPGDGRtFPKRGQtCVVHytG
P62942 T7 Sugiyama FKBP1A FKBP1 FKBP12 ______________MGVQVEtIsPGDGRtFPKRGQtCVVHy
P78345 S253 Sugiyama RPP38 DtsFEDLsKPKRKLADGRQAsVtLQPLKIKKLIPNPNKIRK
P84103 S95 Sugiyama SRSF3 SFRS3 SRP20 RVRVELSNGEKRSRNRGPPPsWGRRPRDDYRRRSPPPRRRS
Q05519 S231 Sugiyama SRSF11 SFRS11 KsDTsSKEIEEAMKRVREAQsLISAAIEPDKKEEKRRHSRS
Q13242 S211 Sugiyama SRSF9 SFRS9 SRP30C GYsRsRSGsRGRDsPyQsRGsPHyFsPFRPy__________
Q13242 S216 Sugiyama SRSF9 SFRS9 SRP30C RSGsRGRDsPyQsRGsPHyFsPFRPy_______________
Q13283 S232 Sugiyama G3BP1 G3BP EEKPEPVLEETAPEDAQKsssPAPADIAQtVQEDLRtFsWA
Q13442 S19 Sugiyama PDAP1 HASPP28 __MPKGGRKGGHKGRARQytsPEEIDAQLQAEKQKAREEEE
Q14152 S1308 Sugiyama EIF3A EIF3S10 KIAA0139 DLRDDRDRRGPPLRSEREEVsSWRRADDRKDDRVEERDPPR
Q14152 S584 Sugiyama EIF3A EIF3S10 KIAA0139 EHQRILARRQtIEERKERLEsLNIQREKEELEQREAELQKV
Q14152 S895 Sugiyama EIF3A EIF3S10 KIAA0139 ERRLGDssLSRKDSRWGDRDsEGtWRKGPEADsEWRRGPPE
Q14155 S518 Sugiyama ARHGEF7 COOL1 KIAA0142 P85SPR PAK3BP PIXB Nbla10314 ERYLLLFPNVLLMLSASPRMsGFIyQGKLPtTGMTITKLED
Q15052 S488 Sugiyama ARHGEF6 COOL2 KIAA0006 PIXA ERYLMLFSNVLIMLSASPRMsGFIYQGKIPIAGTVVTRLDE
Q15185 S72 Sugiyama PTGES3 P23 TEBP EIDLFHCIDPNDsKHKRtDRsILCCLRKGEsGQsWPRLTKE
Q15637 S82 Sugiyama SF1 ZFM1 ZNF162 TRKLRtGDLGIPPNPEDRsPsPEPIyNsEGKRLNTREFRTR
Q16513 S583 Sugiyama PKN2 PRK2 PRKCL2 VTKLDFDLEPEPPPAPPRAssLGEIDEssELRVLDIPGQDs
Q16643 S255 Sugiyama DBN1 D0S117E HRRKQQTLEAEEAKRRLKEQsIFGDHRDEEEEtHMKKsEsE
Q16643 T267 Sugiyama DBN1 D0S117E AKRRLKEQsIFGDHRDEEEEtHMKKsEsEVEEAAAIIAQRP
Q32P51 S95 Sugiyama HNRNPA1L2 HNRNPA1L TPHKVDGRVVEPKRAVsREDsQRPGAHLtVKKIFVGGIKED
Q4G0J3 S285 Sugiyama LARP7 HDCMA18P stEPQKQCSKKKKKRDRVEAssLPEVRTGKRKRsssEDAEs
Q4G0J3 S286 Sugiyama LARP7 HDCMA18P tEPQKQCSKKKKKRDRVEAssLPEVRTGKRKRsssEDAEsL
Q63ZY3 S356 Sugiyama KANK2 ANKRD25 KIAA1518 MXRA3 SIP PREVEVVASTAAGAPAQRAQsLEPYGtGLRALAMPGRPEsP
Q6PCE3 S175 Sugiyama PGM2L1 BM32A PFVPYAVQKLKAVAGVMItAsHNRKEDNGYKVYWETGAQIT
Q6UN15 S492 Sugiyama FIP1L1 FIP1 RHE DRDRDRERERTRERERERDHsPtPsVFNsDEERyRYREYAE
Q6UN15 S496 Sugiyama FIP1L1 FIP1 RHE DRERERTRERERERDHsPtPsVFNsDEERyRYREYAERGYE
Q6UN15 T494 Sugiyama FIP1L1 FIP1 RHE DRDRERERTRERERERDHsPtPsVFNsDEERyRYREYAERG
Q86X29 S493 Sugiyama LSR ILDR3 LISCH QEPAREQAGGGWRARRPRARsVDALDDLtPPstAEsGsRsP
Q8NBJ7 S256 Sugiyama SUMF2 PSEC0171 UNQ1968/PRO4500 EWTASPYQAAEQDMRVLRGAsWIDtADGsANHRARVttRMG
Q8NBS9 S426 Sugiyama TXNDC5 TLP46 UNQ364/PRO700 KKVSEHSGGRDLDSLHRFVLsQAKDEL______________
Q8TAM6 S280 Sugiyama ERMN KIAA1189 TISYRKIRKGNTKQRIDEFEsMMHL________________
Q92995 Y708 GPS6|SIGNOR|PSP USP13 ISOT3 NWIIVHMEEPDFAEPLTMPGyGGAASAGASVFGASGLDNQP
Q96C19 S74 Sugiyama EFHD2 SWS1 ELSAKLLRRADLNQGIGEPQsPsRRVFNPytEFKEFSRKQI
Q96D46 S462 Sugiyama NMD3 CGI-07 FLEDLEEDEAIRKNVNIyRDsAIPVEsDtDDEGAPRISLAE
Q96EP5 S195 Sugiyama DAZAP1 FHDIMGKKVEVKRAEPRDsKsQAPGQPGAsQWGsRVVPNAA
Q96JP5 S83 Sugiyama ZFP91 ZNF757 FKSG11 AAAAAVSRRRKAEYPRRRRssPsARPPDVPGQQPQAAKsPs
Q96MH2 S76 Sugiyama HEXIM2 L3 EDLAGAVGGLGWNsRsPRtQsPGGCsAEAVLARKKHRRRPS
Q96QK1 S786 Sugiyama VPS35 MEM3 TCCCTA00141 INKHFHNtLEHLRLRREsPEsEGPIyEGLIL__________
Q96S44 T8 Sugiyama TP53RK C20orf64 PRPK _____________MAAARAttPADGEEPAPEAEALAAARER
Q96T37 T568 Sugiyama RBM15 OTT OTT1 LVTDAFGHRAPDPLRGARDRtPPLLYRDRDRDLYPDSDWVP
Q9BQE3 S439 Sugiyama TUBA1C TUBA6 SEAREDMAALEKDyEEVGADsADGEDEGEEy__________
Q9BRS2 S507 Sugiyama RIOK1 RIO1 ENQVEERtCsDsEDIGssECsDtDSEEQGDHARPKKHTTDP
Q9BRS2 T509 Sugiyama RIOK1 RIO1 QVEERtCsDsEDIGssECsDtDSEEQGDHARPKKHTTDPDI
Q9BW71 S357 Sugiyama HIRIP3 EDsGKGEPTAKGSRKMARLGstsGEEsDLEREVsDsEAGGG
Q9BXP5 S493 Sugiyama SRRT ARS2 ASR2 VNIKEICWNLQNIRLRECELsPGVNRDLTRRVRNINGITQH
Q9BY44 S524 Sugiyama EIF2A CDA02 MSTP004 MSTP089 DKsPDLAPtPAPQstPRNtVsQsIsGDPEIDKKIKNLKKKL
Q9C0C2 S672 Sugiyama TNKS1BP1 KIAA1741 TAB182 TLARAETtQARtEAQDLCRAsPEPPGPEsSSRWLDDLLAsP
Q9H1E3 S204 Sugiyama NUCKS1 NUCKS JC7 KGKGKVGRPTAsKAsKEKtPsPKEEDEEPEsPPEKKtstsP
Q9H1E3 T202 Sugiyama NUCKS1 NUCKS JC7 PVKGKGKVGRPTAsKAsKEKtPsPKEEDEEPEsPPEKKtst
Q9H7D7 S121 Sugiyama WDR26 CDW2 MIP2 PRO0852 ASSSLATPELGSSLKKKKRLsQsDEDVIRLIGQHLNGLGLN
Q9H7N4 S498 Sugiyama SCAF1 SFRS19 SRA1 GGLDLRRKILTQRRERyRQRsPsPAPAPAPAAAAGPPTRKK
Q9H7N4 S500 Sugiyama SCAF1 SFRS19 SRA1 LDLRRKILTQRRERyRQRsPsPAPAPAPAAAAGPPTRKKSR
Q9NYF8 S196 Sugiyama BCLAF1 BTF KIAA0164 EsPLKsKsQEEPKDtFEHDPsEsIDEFNKssAtsGDIWPGL
Q9NYF8 S319 Sugiyama BCLAF1 BTF KIAA0164 RsPAKtIAPQNAPRDEsRGRssFyPDGGDQETAKtGKFLKR
Q9NYF8 S320 Sugiyama BCLAF1 BTF KIAA0164 sPAKtIAPQNAPRDEsRGRssFyPDGGDQETAKtGKFLKRF
Q9NYV4 S249 Sugiyama CDK12 CRK7 CRKRS KIAA0904 DSSKQDDsPsGAsYGQDyDLsPsRsHtssNYDsYKKsPGsT
Q9NZN8 S242 Sugiyama CNOT2 CDC36 NOT2 HSPC131 MSTP046 VTGLDLSDFPALADRNRREGsGNPtPLINPLAGRAPYVGMV
Q9UI15 S185 Sugiyama TAGLN3 NP25 EQLRQGQNVIGLQMGSNKGAsQAGMtGyGMPRQIM______
Q9UII2 S63 Sugiyama ATP5IF1 ATPI ATPIF1 AGGAFGKREQAEEERyFRAQsREQLAALKKHHEEEIVHHKK
Q9UQ35 S1103 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 sPsLQSKSQTsPKGGRsRsssPVtELAsRsPIRQDRGEFsA
Q9UQ35 S1541 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 ssVDQKtVARtPLGQRsRsGssQELDVKPsAsPQERsEsDs
Q9UQ35 S1542 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 sVDQKtVARtPLGQRsRsGssQELDVKPsAsPQERsEsDss
Q9UQ35 S1648 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 APRALPRRSRSGSSSKGRGPsPEGssstEssPEHPPKSRTA
Q9UQ35 S2132 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 SRMsCFsRPsMsPtPLDRCRsPGMLEPLGssRtPMsVLQQA
Q9UQ35 S2692 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 ERRSRSPRKPIDsLRDsRsLsysPVERRRPsPQPsPRDQQS
Q9UQ35 S323 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 RGEGDAPFsEPGttstQRPssPEtAtKQPssPYEDKDKDKK
Q9UQ35 S351 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 PssPYEDKDKDKKEKsAtRPsPsPERsstGPEPPAPtPLLA
Q9UQ35 S353 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 sPYEDKDKDKKEKsAtRPsPsPERsstGPEPPAPtPLLAER
Q9UQ35 S358 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 KDKDKKEKsAtRPsPsPERsstGPEPPAPtPLLAERHGGsP
Q9UQ35 T359 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 DKDKKEKsAtRPsPsPERsstGPEPPAPtPLLAERHGGsPQ
Q9Y2B0 S115 Sugiyama CNPY2 MSAP TMEM4 ZSIG9 UNQ1943/PRO4426 QIDPstHRKNYVRVVGRNGEssELDLQGIRIDsDISGtLKF
Q9Y3U8 S91 Sugiyama RPL36 KFIKKRVGTHIRAKRKREELsNVLAAMRKAAAKKD______
Q9Y5S9 S56 Sugiyama RBM8A RBM8 HSPC114 MDS014 KGRGFGsEEGsRARMREDyDsVEQDGDEPGPQRSVEGWILF