HASPIN

All methods over the ±5 window, side by side.
PSPA
CDDM
MLP-attr
Normalized values from positional scanning peptide array
PSPA measures pT, so at flanking positions the pS row is a duplicate of the pT row. At position 0 the acceptor identity (pS, pT, pY) is genuine.
Position-wise Probabilities
Log-Odds: Probabilities / STY Background
Position-wise Probabilities (±5 window)
Log-Odds (±5 window)
Motifs stratified by phospho-acceptor (S / T / Y)
K-means sub-motif clusters
Signed attribution PSSM distilled from the CDDM discriminative MLP (in-silico saturation mutagenesis). Letters above 0 are enriched, below are depleted; this is not a probability distribution.
MLP-attribution PSSM (±5, signed)
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substrate_uniprot site source substrate_genes site_seq
O15371 T49 Sugiyama EIF3D EIF3S7 PyQPFsKGDRLGKVADWtGAtyQDKRYTNKYSSQFGGGSQY
O43390 T137 Sugiyama HNRNPR HNRPR ESTKGPDEAKIKALLERtGytLDVttGQRKYGGPPPDSVYS
O60506 T134 Sugiyama SYNCRIP HNRPQ NSAP1 DSSKGPDEAKIKALLERtGytLDVttGQRKYGGPPPDSVYS
O60841 S186 Sugiyama EIF5B IF2 KIAA0741 EDEDNsKKIKERsRINssGEsGDEsDEFLQsRKGQKKNQKN
O95218 S181 Sugiyama ZRANB2 ZIS ZNF265 DEDLsKyKLDEDEDEDDADLsKyNLDAsEEEDsNKKKSNRR
P00558 T168 Sugiyama PGK1 PGKA MIG10 OK/SW-cl.110 EAFRAsLSKLGDVyVNDAFGtAHRAHssMVGVNLPQKAGGF
P02545 S583 Sugiyama LMNA LMN1 HHHHGsHCsssGDPAEYNLRsRtVLCGtCGQPADKAsAsGs
P04792 S82 Sugiyama HSPB1 HSP27 HSP28 AIEsPAVAAPAYsRALsRQLssGVsEIRHtADRWRVsLDVN
P07205 T168 Sugiyama PGK2 PGKB EAFRAsLSKLGDVyVNDAFGtAHRAHSSMVGVNLPHKASGF
P10412 S55 Sugiyama H1-4 H1F4 HIST1H1E AsGPPVsELItKAVAASKERsGVsLAALKKALAAAGyDVEK
P13639 S593 Sugiyama EEF2 EF2 SDPVVsyREtVsEEsNVLCLsKsPNKHNRLyMKARPFPDGL
P15880 T202 Sugiyama RPS2 RPS4 CKVTGRCGsVLVRLIPAPRGtGIVsAPVPKKLLMMAGIDDC
P15880 T232 Sugiyama RPS2 RPS4 KLLMMAGIDDCytsARGCtAtLGNFAKATFDAISKTysyLt
P15880 T278 Sugiyama RPS2 RPS4 EtVFtKsPyQEFtDHLVKtHtRVsVQRtQAPAVATt_____
P15880 T285 Sugiyama RPS2 RPS4 PyQEFtDHLVKtHtRVsVQRtQAPAVATt____________
P16402 S56 Sugiyama H1-3 H1F3 HIST1H1D AsGPPVsELItKAVAASKERsGVsLAALKKALAAAGyDVEK
P16403 S55 Sugiyama H1-2 H1F2 HIST1H1C AsGPPVsELItKAVAASKERsGVsLAALKKALAAAGyDVEK
P17812 S574 Sugiyama CTPS1 CTPS YLQKGCRLsPRDtysDRsGsssPDsEItELKFPsINHD___
P18077 Y14 Sugiyama RPL35A GIG33 _______MSGRLWSKAIFAGyKRGLRNQREHTALLKIEGVy
P18124 T17 Sugiyama RPL7 ____MEGVEEKKKEVPAVPEtLKKKRRNFAELKIKRLRKKF
P18583 S2011 Sugiyama SON C21orf50 DBP5 KIAA1019 NREBP HSPC310 HSPC312 RSRTPSRRRRSRSVVRRRsFsIsPVRLRRSRTPLRRRFsRs
P18621 S5 Sugiyama RPL17 ________________MVRysLDPENPtKsCKSRGSNLRVH
P19338 S458 Sugiyama NCL TEADAEKTFEEKQGtEIDGRsIsLyytGEKGQNQDYRGGKN
P20618 S157 Sugiyama PSMB1 PSC5 IIGGLDEEGKGAVysFDPVGsyQRDsFKAGGSASAMLQPLL
P22090 S204 Sugiyama RPS4Y1 RPS4Y PRO2646 IGGANLGRVGVITNRERHPGsFDVVHVKDANGNSFATRLSN
P22626 S351 Sugiyama HNRNPA2B1 HNRPA2B1 yGGGNyGPGGsGGsGGyGGRsRy__________________
P22626 Y347 Sugiyama HNRNPA2B1 HNRPA2B1 MGGPyGGGNyGPGGsGGsGGyGGRsRy______________
P26373 S77 Sugiyama RPL13 BBC1 OK/SW-cl.46 PIVRCPTVRYHTKVRAGRGFsLEELRVAGIHKKVARTIGIs
P35268 T62 Sugiyama RPL22 FLQERIKVNGKAGNLGGGVVtIERsKSKITVTSEVPFSKRY
P36578 S87 Sugiyama RPL4 RPL1 WGtGRAVARIPRVRGGGtHRsGQGAFGNMCRGGRMFAPTKT
P36578 T69 Sugiyama RPL4 RPL1 RQPyAVsELAGHQtsAEsWGtGRAVARIPRVRGGGtHRsGQ
P36578 T84 Sugiyama RPL4 RPL1 AEsWGtGRAVARIPRVRGGGtHRsGQGAFGNMCRGGRMFAP
P37108 T45 Sugiyama SRP14 sVyItLKKYDGRTKPIPKKGtVEGFEPADNKCLLRATDGKK
P38646 T120 Sugiyama HSPA9 GRP75 HSPA9B mt-HSP70 LVGMPAKRQAVtNPNNtFyAtKRLIGRRYDDPEVQKDIKNV
P38646 T86 Sugiyama HSPA9 GRP75 HSPA9B mt-HSP70 CVAVMEGKQAKVLENAEGARttPsVVAFtADGERLVGMPAK
P39023 S13 Sugiyama RPL3 OK/SW-cl.32 ________MSHRKFSAPRHGsLGFLPRKRSSRHRGKVKSFP
P42677 S78 Sugiyama RPS27 MPS1 GCSTVLCQPTGGKARLtEGCsFRRKQH______________
P46778 T124 Sugiyama RPL21 sFLKRVKENDQKKKEAKEKGtWVQLKRQPAPPREAHFVRTN
P46778 T29 Sugiyama RPL21 RGTRYMFSRPFRKHGVVPLAtyMRIYKKGDIVDIKGMGTVQ
P46779 T81 Sugiyama RPL28 VVVIKRRSGQRKPAtsyVRttINKNARAtLsSIRHMIRKNK
P46783 T148 Sugiyama RPS10 RRSAVPPGADKKAEAGAGsAtEFQFRGGFGRGRGQPPQ___
P47914 T151 Sugiyama RPL29 DQTKAQAAAPAsVPAQAPKRtQAPtKAsE____________
P47914 T155 Sugiyama RPL29 AQAAAPAsVPAQAPKRtQAPtKAsE________________
P50914 T43 Sugiyama RPL14 KLVAIVDVIDQNRALVDGPCtQVRRQAMPFKCMQLTDFILK
P60709 S323 Sugiyama ACTB ttMyPGIADRMQKEItALAPstMKIKIIAPPERKYsVWIGG
P60709 T324 Sugiyama ACTB tMyPGIADRMQKEItALAPstMKIKIIAPPERKYsVWIGGs
P61313 S97 Sugiyama RPL15 EC45 TCBAP0781 KGAtyGKPVHHGVNQLKFARsLQsVAEERAGRHCGALRVLN
P61313 Y6 Sugiyama RPL15 EC45 TCBAP0781 _______________MGAYKyIQELWRKKQSDVMRFLLRVR
P61353 S39 Sugiyama RPL27 SGRKAVIVKNIDDGtsDRPysHALVAGIDRyPRKVTAAMGK
P61513 T16 Sugiyama RPL37A _____MAKRTKKVGIVGKyGtRYGAsLRKMVKKIEISQHAK
P62081 T121 Sugiyama RPS7 LPKPTRKSRTKNKQKRPRsRtLtAVHDAILEDLVFPsEIVG
P62081 T123 Sugiyama RPS7 KPTRKSRTKNKQKRPRsRtLtAVHDAILEDLVFPsEIVGKR
P62249 S9 Sugiyama RPS16 ____________MPsKGPLQsVQVFGRKKTATAVAHCKRGN
P62263 S139 Sugiyama RPS14 PRO2640 LARSGMKIGRIEDVtPIPsDstRRKGGRRGRRL________
P62263 T140 Sugiyama RPS14 PRO2640 ARSGMKIGRIEDVtPIPsDstRRKGGRRGRRL_________
P62273 Y7 Sugiyama RPS29 ______________MGHQQLyWsHPRKFGQGSRSCRVCSNR
P62277 S12 Sugiyama RPS13 _________MGRMHAPGKGLsQsALPyRRsVPtWLKLTsDD
P62277 S14 Sugiyama RPS13 _______MGRMHAPGKGLsQsALPyRRsVPtWLKLTsDDVK
P62280 T15 Sugiyama RPS11 ______MADIQTERAyQKQPtIFQNKKRVLLGEtGKEKLPR
P62701 S204 Sugiyama RPS4X CCG2 RPS4 SCAR tGGANLGRIGVITNRERHPGsFDVVHVKDANGNSFATRLsN
P62736 S325 Sugiyama ACTA2 ACTSA ACTVS GIG46 tTMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGG
P62736 T326 Sugiyama ACTA2 ACTSA ACTVS GIG46 TMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGGS
P62750 T42 Sugiyama RPL23A LKAKKAVLKGVHSHKKKKIRtsPTFRRPKTLRLRRQPKYPR
P62899 S15 Sugiyama RPL31 ______MAPAKKGGEKKKGRsAINEVVtREytINIHKRIHG
P62899 T26 Sugiyama RPL31 KGGEKKKGRsAINEVVtREytINIHKRIHGVGFKKRAPRAL
P62910 S105 Sugiyama RPL32 PP9932 LEVLLMCNKsyCAEIAHNVSsKNRKAIVERAAQLAIRVTNP
P62917 S14 Sugiyama RPL8 _______MGRVIRGQRKGAGsVFRAHVKHRKGAARLRAVDF
P62917 T135 Sugiyama RPL8 CLEEKPGDRGKLARAsGNyAtVIsHNPEtKKtRVKLPSGSK
P62917 T143 Sugiyama RPL8 RGKLARAsGNyAtVIsHNPEtKKtRVKLPSGSKKVISSANR
P62917 T146 Sugiyama RPL8 LARAsGNyAtVIsHNPEtKKtRVKLPSGSKKVISSANRAVV
P63261 S323 Sugiyama ACTG1 ACTG ttMyPGIADRMQKEItALAPstMKIKIIAPPERKYsVWIGG
P63261 T324 Sugiyama ACTG1 ACTG tMyPGIADRMQKEItALAPstMKIKIIAPPERKYsVWIGGs
P63267 S324 Sugiyama ACTG2 ACTA3 ACTL3 ACTSG tTMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGG
P63267 T325 Sugiyama ACTG2 ACTA3 ACTL3 ACTSG TMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGGS
P67809 S136 Sugiyama YBX1 NSEP1 YB1 GEKGAEAANVtGPGGVPVQGsKYAADRNHYRRYPRRRGPPR
P67809 Y196 Sugiyama YBX1 NSEP1 YB1 sAPEGQAQQRRPyRRRRFPPyyMRRPYGRRPQysNPPVQGE
P68032 S325 Sugiyama ACTC1 ACTC tTMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGG
P68032 T326 Sugiyama ACTC1 ACTC TMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGGS
P68133 S325 Sugiyama ACTA1 ACTA TTMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGG
P68133 T326 Sugiyama ACTA1 ACTA TMYPGIADRMQKEItALAPstMKIKIIAPPERKYSVWIGGS
P68431 T4 GPS6|SIGNOR|EPSD H3C1 H3FA HIST1H3A; H3C2 H3FL HIST1H3B; H3C3 H3FC HIST1H3C; H3C4 H3FB HIST1H3D; H3C6 H3FD HIST1H3E; H3C7 H3FI HIST1H3F; H3C8 H3FH HIST1H3G; H3C10 H3FK HIST1H3H; H3C11 H3FF HIST1H3I; H3C12 H3FJ HIST1H3J _________________MARtKQtARKstGGKAPRKQLATK
P83731 T52 Sugiyama RPL24 LNAKCEsAFLSKRNPRQINWtVLyRRKHKKGQsEEIQKKRT
P83731 Y55 Sugiyama RPL24 KCEsAFLSKRNPRQINWtVLyRRKHKKGQsEEIQKKRTRRA
P84098 S37 Sugiyama RPL19 CGKKKVWLDPNEtNEIANANsRQQIRKLIKDGLIIRKPVtV
P84243 T4 GPS6|EPSD H3-3A H3.3A H3F3 H3F3A PP781; H3-3B H3.3B H3F3B _________________MARtKQTARKstGGKAPRKQLATK
Q00059 S160 Sugiyama TFAM TCF6 TCF6L2 RKAMTKKKELTLLGKPKRPRsAYNVyVAERFQEAKGDsPQE
Q01105 S168 Sugiyama SET ENKVLSKEFHLNEsGDPssKstEIKWKSGKDLTKRssQTQN
Q02543 S123 Sugiyama RPL18A AGAVtQCyRDMGARHRARAHsIQIMKVEEIAASKCRRPAVK
Q04695 T410 Sugiyama KRT17 GEDAHLTQyKKEPVTTRQVRtIVEEVQDGKVISSREQVHQT
Q07020 T136 Sugiyama RPL18 AGGKILtFDQLALDsPKGCGtVLLsGPRKGREVYRHFGKAP
Q07020 T85 Sugiyama RPL18 RMIRKMKLPGRENKtAVVVGtItDDVRVQEVPKLKVCALRV
Q13263 S471 Sugiyama TRIM28 KAP1 RNF96 TIF1B FGsGDDPyssAEPHVsGVKRsRsGEGEVsGLMRKVPRVsLE
Q13442 S63 Sugiyama PDAP1 HASPP28 GGDGAAGDPKKEKKsLDsDEsEDEEDDyQQKRKGVEGLIDI
Q16181 T426 Sugiyama SEPTIN7 CDC10 SEPT7 DEKANWEAQQRILEQQNssRtLEKNKKKGKIF_________
Q16695 T4 GPS6 H3-4 H3FT HIST3H3 _________________MARtKQTARKsTGGKAPRKQLATK
Q71DI3 T4 GPS6|EPSD H3C15 HIST2H3A; H3C14 H3F2 H3FM HIST2H3C; H3C13 HIST2H3D _________________MARtKQTARKsTGGKAPRKQLATK
Q71UM5 S78 Sugiyama RPS27L GCSTVLCQPTGGKARLtEGCsFRRKQH______________
Q86V81 S145 Sugiyama ALYREF ALY BEF THOC4 LFAEFGtLKKAAVHYDRsGRsLGtADVHFERKADALKAMKQ
Q8TF76 S108 Sugiyama HASPIN GSG2 PKDRPsLTVtPKRWKLRARPsLtVtPRRLGLRARPPQKCSt
Q8TF76 S147 Sugiyama HASPIN GSG2 StPCGPLRLPPFPsRDsGRLsPDLsVCGQPRDGDELGIsAS
Q8TF76 S151 Sugiyama HASPIN GSG2 GPLRLPPFPsRDsGRLsPDLsVCGQPRDGDELGIsASLFSS
Q8TF76 S288 Sugiyama HASPIN GSG2 EsCCKRKLVVGNGPEGPGLsstGKRRAtGQDSCQERGLQEA
Q8TF76 S349 Sugiyama HASPIN GSG2 DRLERTRSSRKSKHQEATEtsLLHsHRFKKGQKLGKDsFPT
Q8TF76 S353 Sugiyama HASPIN GSG2 RTRSSRKSKHQEATEtsLLHsHRFKKGQKLGKDsFPTQDLt
Q8TF76 S389 Sugiyama HASPIN GSG2 TQDLtPLQNVCFWTKTRAsFsFHKKKIVTDVSEVCSIYTTA
Q8TF76 S543 Sugiyama HASPIN GSG2 GSHQKTFEEILPEIIISKELsLLSGEVCNRTEGFIGLNSVH
Q8TF76 T128 Sugiyama HASPIN GSG2 sLtVtPRRLGLRARPPQKCStPCGPLRLPPFPsRDsGRLsP
Q8TF76 T15 Sugiyama HASPIN GSG2 ______MAAsLPGPGsRLFRtyGAADGRRQRRPGREAAQWF
Q8TF76 T289 Sugiyama HASPIN GSG2 sCCKRKLVVGNGPEGPGLsstGKRRAtGQDSCQERGLQEAV
Q8TF76 Y16 Sugiyama HASPIN GSG2 _____MAAsLPGPGsRLFRtyGAADGRRQRRPGREAAQWFP
Q92522 T55 Sugiyama H1-10 H1FX KRKNSKKKNQPGKySQLVVEtIRRLGERNGSSLAKIYTEAK
Q92882 T200 Sugiyama OSTF1 ATNAACAsLLKKKQGTDAVRtLsNAEDyLDDEDsD______
Q99543 T268 Sugiyama DNAJC2 MPHOSPH11 MPP11 ZRF1 EKQNRATRAQRKKEEMNRIRtLVDNAysCDPRIKKFKEEEK
Q99613 T880 Sugiyama EIF3C EIF3S8 AEKLGsLVENNERVFDHKQGtyGGyFRDQKDGYRKNEGYMR
Q9BQ69 T196 Sugiyama MACROD1 LRP16 SLLGGGGVDGCIHRAAGPLLtDECRtLQSCKTGKAKITGGY
Q9BQ69 T201 Sugiyama MACROD1 LRP16 GGVDGCIHRAAGPLLtDECRtLQSCKTGKAKITGGYRLPAK
Q9H0H5 S206 Sugiyama RACGAP1 KIAA1478 MGCRACGAP tsRQFVDGPPGPVKKtRsIGsAVDQGNEsIVAKTTVTVPND
Q9H0H5 T251 Sugiyama RACGAP1 KIAA1478 MGCRACGAP EAVSTIETVPYWtRSRRKtGtLQPWNsDstLNSRQLEPRTE
Q9NQ39 T159 Sugiyama RPS10P5 RPS10L RRCSVPPGADKKAEAGAGsAtEFQFRGRCGRGRGQPPQ___
Q9NZB2 T991 Sugiyama FAM120A C9orf10 KIAA0183 OSSA PLQVVSVGGPARGRPRGVIstPVIRTFGRGGRYYGRGYKNQ
Q9P258 T521 Sugiyama RCC2 KIAA1470 TD60 RDESETEKEKIKKLPEYNPRtL___________________
Q9UGV2 S327 Sugiyama NDRG3 KYFLQGMGYIPSASMTRLARsRtHstsssLGsGEsPFsRsV
Q9UGV2 T329 Sugiyama NDRG3 FLQGMGYIPSASMTRLARsRtHstsssLGsGEsPFsRsVts
Q9UQ35 S1541 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 ssVDQKtVARtPLGQRsRsGssQELDVKPsAsPQERsEsDs
Q9UQ35 S2102 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 GSSSDRSRsAtPPATRNHsGsRtPPVALNsSRMsCFsRPsM
Q9Y2V2 T50 Sugiyama CARHSP1 sPsPLRGNVVPsPLPtRRtRtFsAtVRASQGPVYKGVCKCF
Q9Y2X3 S440 Sugiyama NOP58 NOL5 NOP5 HSPC120 EHKSEVKtyDPSGDstLPtCsKKRKIEQVDKEDEITEKKAK
Q9Y388 S227 Sugiyama RBMX2 CGI-79 SKNSERAQKSEPREGQKLPKsRTAYsGGAEDLERELKKEKP
Q9Y3U8 Y53 Sugiyama RPL36 LTKHTKFVRDMIREVCGFAPyERRAMELLKVSKDKRALKFI