HIPK1

All methods over the ±5 window, side by side.
PSPA
CDDM
MLP-attr
Normalized values from positional scanning peptide array
PSPA measures pT, so at flanking positions the pS row is a duplicate of the pT row. At position 0 the acceptor identity (pS, pT, pY) is genuine.
Position-wise Probabilities
Log-Odds: Probabilities / STY Background
Position-wise Probabilities (±5 window)
Log-Odds (±5 window)
Motifs stratified by phospho-acceptor (S / T / Y)
K-means sub-motif clusters
Signed attribution PSSM distilled from the CDDM discriminative MLP (in-silico saturation mutagenesis). Letters above 0 are enriched, below are depleted; this is not a probability distribution.
MLP-attribution PSSM (±5, signed)
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substrate_uniprot site source substrate_genes site_seq
O00115 S70 Sugiyama DNASE2 DNASE2A DNL2 QYKyLDEssGGWRDGRALINsPEGAVGRSLQPLyRSNTSQL
O00165 S153 Sugiyama HAX1 HS1BP1 PDSHQPRIFGGVLESDARSEsPQPAPDWGsQRPFHRFDDVW
O14639 S431 Sugiyama ABLIM1 ABLIM KIAA0059 LIMAB1 TSGYDDKQERQsLGEsPRtLsPtPsAEGyQDVRDRMIHRst
O60716 S252 Sugiyama CTNND1 KIAA0384 sRVtRIEERyRPsMEGyRAPsRQDVyGPQPQVRVGGssVDL
O60829 T51 SIGNOR|EPSD|PSP PAGE4 GAGEC1 JM27 QQEEPPTDNQDIEPGQEREGtPPIEERKVEGDCQEMDLEKt
P04075 S46 Sugiyama ALDOA ALDA PGKGILAADEstGsIAKRLQsIGtENtEENRRFyRQLLLtA
P04406 S241 Sugiyama GAPDH GAPD CDABP0047 OK/SW-cl.12 IPELNGKLtGMAFRVPtANVsVVDLtCRLEKPAKyDDIKKV
P04406 T237 Sugiyama GAPDH GAPD CDABP0047 OK/SW-cl.12 VGKVIPELNGKLtGMAFRVPtANVsVVDLtCRLEKPAKyDD
P04792 S78 Sugiyama HSPB1 HSP27 HSP28 LPPAAIEsPAVAAPAYsRALsRQLssGVsEIRHtADRWRVs
P04792 S82 Sugiyama HSPB1 HSP27 HSP28 AIEsPAVAAPAYsRALsRQLssGVsEIRHtADRWRVsLDVN
P07195 S303 Sugiyama LDHB GIENEVFLSLPCILNARGLtsVINQKLKDDEVAQLKKsADt
P07737 S30 Sugiyama PFN1 NLMADGTCQDAAIVGYKDsPsVWAAVPGKtFVNITPAEVGV
P08195 S607 Sugiyama SLC3A2 MDU1 ASLPAKADLLLstQPGREEGsPLELERLKLEPHEGLLLRFP
P10586 S1853 Sugiyama PTPRF LAR GPITVHCSAGVGRtGVFItLsIVLERMRyEGVVDMFQTVKT
P10586 T1846 Sugiyama PTPRF LAR KEQFGQDGPITVHCSAGVGRtGVFItLsIVLERMRyEGVVD
P10586 T1851 Sugiyama PTPRF LAR QDGPITVHCSAGVGRtGVFItLsIVLERMRyEGVVDMFQTV
P10809 S70 Sugiyama HSPD1 HSP60 AVAVTMGPKGRtVIIEQsWGsPKVTKDGVtVAKsIDLKDKY
P11142 T37 Sugiyama HSPA8 HSC70 HSP73 HSPA10 CVGVFQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKN
P11142 T38 Sugiyama HSPA8 HSC70 HSP73 HSPA10 VGVFQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKNQ
P11142 Y41 Sugiyama HSPA8 HSC70 HSP73 HSPA10 FQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKNQVAM
P17812 S573 Sugiyama CTPS1 CTPS HYLQKGCRLsPRDtysDRsGsssPDsEItELKFPsINHD__
P17812 S575 Sugiyama CTPS1 CTPS LQKGCRLsPRDtysDRsGsssPDsEItELKFPsINHD____
P22234 S27 Sugiyama PAICS ADE2 AIRC PAIS LNIGKKLYEGKtKEVyELLDsPGKVLLQsKDQItAGNAARK
P22626 S189 Sugiyama HNRNPA2B1 HNRPA2B1 IVLQKyHtINGHNAEVRKALsRQEMQEVQssRsGRGGNFGF
P23468 S1858 Sugiyama PTPRD GPISVHCSAGVGRtGVFItLsIVLERMRYEGVVDIFQTVKM
P23468 T1851 Sugiyama PTPRD KEQFGQDGPISVHCSAGVGRtGVFItLsIVLERMRYEGVVD
P23468 T1856 Sugiyama PTPRD QDGPISVHCSAGVGRtGVFItLsIVLERMRYEGVVDIFQTV
P25054 S1389 Sugiyama APC DP2.5 PKsPPEHYVQETPLMFsRCTsVSsLDSFESRSIASSVQSEP
P27348 S45 Sugiyama YWHAQ CMKAVtEQGAELsNEERNLLsVAyKNVVGGRRSAWRVIsSI
P27824 S362 Sugiyama CANX WDEDMDGEWEAPQIANPRCEsAPGCGVWQRPVIDNPNyKGK
P29401 S295 Sugiyama TKT ysQIQsKKKILAtPPQEDAPsVDIANIRMPsLPsyKVGDKI
P29401 T287 Sugiyama TKT AEQIIQEIysQIQsKKKILAtPPQEDAPsVDIANIRMPsLP
P29692 S162 Sugiyama EEF1D EF1D AKKPAtPAEDDEDDDIDLFGsDNEEEDKEAAQLREERLRQY
P29692 T147 Sugiyama EEF1D EF1D PQtQHVsPMRQVEPPAKKPAtPAEDDEDDDIDLFGsDNEEE
P30086 S52 Sugiyama PEBP1 PBP PEBP AAVDELGKVLtPtQVKNRPtsIsWDGLDsGKLytLVLtDPD
P30086 T51 Sugiyama PEBP1 PBP PEBP GAAVDELGKVLtPtQVKNRPtsIsWDGLDsGKLytLVLtDP
P31946 S47 Sugiyama YWHAB AMKAVtEQGHELsNEERNLLsVAyKNVVGARRSsWRVIsSI
P31947 S45 Sugiyama SFN HME1 FMKGAVEKGEELsCEERNLLsVAyKNVVGGQRAAWRVLSsI
P34931 T39 Sugiyama HSPA1L CVGVFQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKN
P34931 T40 Sugiyama HSPA1L VGVFQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKNQ
P34931 Y43 Sugiyama HSPA1L FQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKNQVAM
P35269 S385 Sugiyama GTF2F1 RAP74 KKKTPPKRERKPsGGSSRGNsRPGtPsAEGGstsstLRAAA
P35269 T389 Sugiyama GTF2F1 RAP74 PPKRERKPsGGSSRGNsRPGtPsAEGGstsstLRAAASKLE
P38646 S89 Sugiyama HSPA9 GRP75 HSPA9B mt-HSP70 VMEGKQAKVLENAEGARttPsVVAFtADGERLVGMPAKRQA
P38646 T87 Sugiyama HSPA9 GRP75 HSPA9B mt-HSP70 VAVMEGKQAKVLENAEGARttPsVVAFtADGERLVGMPAKR
P42166 T160 Sugiyama TMPO LAP2 KLYEKKLLKLREQGtEsRsstPLPtIsssAENtRQNGsNDs
P42167 T160 Sugiyama TMPO LAP2 KLYEKKLLKLREQGTESRsstPLPTISsSAENTRQNGSNDS
P51608 S216 EPSD|PSP MECP2 TTRPKAATSEGVQVKRVLEKsPGKLLVKMPFQtsPGGKAEG
P51608 S80 EPSD|PSP MECP2 GKAETsEGsGsAPAVPEAsAsPKQRRSIIRDRGPMYDDPTL
P54652 T38 Sugiyama HSPA2 CVGVFQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKN
P54652 T39 Sugiyama HSPA2 VGVFQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKNQ
P54652 Y42 Sugiyama HSPA2 FQHGKVEIIANDQGNRttPsyVAFtDtERLIGDAAKNQVAM
P60174 S212 Sugiyama TPI1 TPI WLKsNVsDAVAQstRIIyGGsVtGAtCKELASQPDVDGFLV
P61981 S46 Sugiyama YWHAG AMKNVtELNEPLsNEERNLLsVAyKNVVGARRSsWRVIsSI
P62258 S46 Sugiyama YWHAE SMKKVAGMDVELTVEERNLLsVAyKNVIGARRASWRIIssI
P62263 T133 Sugiyama RPS14 PRO2640 QsALRALARSGMKIGRIEDVtPIPsDstRRKGGRRGRRL__
P63104 S45 Sugiyama YWHAZ CMKsVtEQGAELsNEERNLLsVAyKNVVGARRssWRVVssI
Q00613 S363 Sugiyama HSF1 HSTF1 AsVTALtDARGHTDtEGRPPsPPPtstPEKCLsVACLDKNE
Q04917 S46 Sugiyama YWHAH YWHA1 AMKAVtELNEPLsNEDRNLLsVAyKNVVGARRSsWRVIsSI
Q07666 S35 Sugiyama KHDRBS1 SAM68 sGRsGsMDPsGAHPsVRQtPsRQPPLPHRSRGGGGGSRGGA
Q13283 S231 Sugiyama G3BP1 G3BP QEEKPEPVLEETAPEDAQKsssPAPADIAQtVQEDLRtFsW
Q13283 S232 Sugiyama G3BP1 G3BP EEKPEPVLEETAPEDAQKsssPAPADIAQtVQEDLRtFsWA
Q13332 S1894 Sugiyama PTPRS GPISVHCSAGVGRtGVFItLsIVLERMRYEGVVDIFQTVKM
Q13332 T1887 Sugiyama PTPRS KEQFGQDGPISVHCSAGVGRtGVFItLsIVLERMRYEGVVD
Q13332 T1892 Sugiyama PTPRS QDGPISVHCSAGVGRtGVFItLsIVLERMRYEGVVDIFQTV
Q13428 T1014 Sugiyama TCOF1 ARssssEsEDEDVIPAtQCLtPGIRTNVVTMPTAHPRIAPK
Q13573 S182 Sugiyama SNW1 SKIIP SKIP AMPVRAADKLAPAQyIRytPsQQGVAFNsGAKQRVIRMVEM
Q13586 S620 Sugiyama STIM1 GOK sPsAPPGGsPHLDssRsHsPssPDPDtPsPVGDSRALQASR
Q14160 S1448 Sugiyama SCRIB CRIB1 KIAA0147 LAP4 SCRB1 VARTUL QEDEQPPWAsPsPtsRQsPAsPPPLGGGAPVRTAKAERRHQ
Q14247 S405 Sugiyama CTTN EMS1 EEARRKLEEQARAKtQtPPVsPAPQPtEERLPssPVyEDAA
Q14258 S100 Sugiyama TRIM25 EFP RNF147 ZNF147 DLAREPPADVWtPPARAsAPsPNAQVACDHCLKEAAVKTCL
Q15293 T76 Sugiyama RCN1 RCN FQYDHEAFLGKEDsKtFDQLtPDEsKERLGKIVDRIDNDGD
Q15637 S82 Sugiyama SF1 ZFM1 ZNF162 TRKLRtGDLGIPPNPEDRsPsPEPIyNsEGKRLNTREFRTR
Q71RC2 S597 Sugiyama LARP4 PP13296 QttIPVsPPsttKPsRAstAsPCNNNINAAtAVALQEPRKL
Q7Z2W4 T273 Sugiyama ZC3HAV1 ZC3HDC2 PRO1677 FFQGsQEFLASASAsAERsCtPsPDQIsHRAsLEDAPVDDL
Q86YP4 S100 Sugiyama GATAD2A DGPVDMRTSHSDMKSERRPPsPDVIVLsDNEQPssPRVNGL
Q86Z02 S239 Sugiyama HIPK1 KIAA0630 MYAK NBAK2 KILKNHPSYARQGQIEVSILsRLSsENADEYNFVRSYECFQ
Q86Z02 S243 Sugiyama HIPK1 KIAA0630 MYAK NBAK2 NHPSYARQGQIEVSILsRLSsENADEYNFVRSYECFQHKNH
Q86Z02 S355 Sugiyama HIPK1 KIAA0630 MYAK NBAK2 VIDFGSAsHVSKAVCstyLQsRYYRAPEIILGLPFCEAIDM
Q86Z02 Y352 Sugiyama HIPK1 KIAA0630 MYAK NBAK2 RVKVIDFGSAsHVSKAVCstyLQsRYYRAPEIILGLPFCEA
Q8IWW6 S240 Sugiyama ARHGAP12 SSSsTEQIRAttPPNQGRPDsPVyANLQELKISQSALPPLP
Q8IWW6 T231 Sugiyama ARHGAP12 QDSEsGDELSSSsTEQIRAttPPNQGRPDsPVyANLQELKI
Q8IWX8 S817 Sugiyama CHERP DAN26 SCAF6 QSRSRSKSYSPGRRRRsRsRsPtPPssAGLGsNsAPPIPDS
Q8IWX8 T819 Sugiyama CHERP DAN26 SCAF6 RSRSKSYSPGRRRRsRsRsPtPPssAGLGsNsAPPIPDSRL
Q8IZ21 S118 Sugiyama PHACTR4 PRO2963 PSDAMLKNGHTTPIGNARsssPVQVEEEPVRLAsLRKAIPE
Q8N684 T203 Sugiyama CPSF7 GGIPPRAHsRDssDsADGRAtPsENLVPSSARVDKPPSVLP
Q8ND56 S183 Sugiyama LSM14A C19orf13 FAM61A RAP55 RAP55A GSAFTQDTRsLKtQLsQGRssPQLDPLRKsPtMEQAVQtAs
Q8NFD5 S1744 Sugiyama ARID1B BAF250B DAN15 KIAA1235 OSA2 ALDTINILLYDDSTVATFNLsQLSGFLELLVEYFRKCLIDI
Q8TEW0 S962 Sugiyama PARD3 PAR3 PAR3A DEGMEtLEEDTEESSRsGREsVstAsDQPsHsLERQMNGNQ
Q8WXI9 S122 Sugiyama GATAD2B KIAA1150 DEPVDMSARRsEPERGRLtPsPDIIVLsDNEAssPRsSsRM
Q8WXI9 T120 Sugiyama GATAD2B KIAA1150 INDEPVDMSARRsEPERGRLtPsPDIIVLsDNEAssPRsSs
Q8WY36 S844 Sugiyama BBX HBP2 KRKARKTKITHLVRtADGRVsPAGGTLDDKPKEQLQRSLPK
Q96B36 S212 Sugiyama AKT1S1 PRAS40 GFKEKRtEARssDEENGPPssPDLDRIAAsMRALVLREAED
Q96G74 S64 Sugiyama OTUD5 GVGGGDRDRDsGVVGARPRAsPPPQGPLPGPPGALHRWALA
Q96MH2 S76 Sugiyama HEXIM2 L3 EDLAGAVGGLGWNsRsPRtQsPGGCsAEAVLARKKHRRRPS
Q96MH2 T74 Sugiyama HEXIM2 L3 EDEDLAGAVGGLGWNsRsPRtQsPGGCsAEAVLARKKHRRR
Q9C0C2 S1103 Sugiyama TNKS1BP1 KIAA1741 TAB182 KRGWVGEFSLSVGPQREAAFsPGQQDWsRDFCIEASERsyQ
Q9C0C2 S672 Sugiyama TNKS1BP1 KIAA1741 TAB182 TLARAETtQARtEAQDLCRAsPEPPGPEsSSRWLDDLLAsP
Q9H2X6 S364 Sugiyama HIPK2 VIDFGSAsHVSKAVCstyLQsRYyRAPEIILGLPFCEAIDM
Q9H2X6 Y361 Sugiyama HIPK2 RVKVIDFGSAsHVSKAVCstyLQsRYyRAPEIILGLPFCEA
Q9HC35 S144 Sugiyama EML4 C2orf2 EMAPL4 GQREKKEEsHsNDQsPQIRAsPsPQPssQPLQIHRQtPEsK
Q9HC35 S146 Sugiyama EML4 C2orf2 EMAPL4 REKKEEsHsNDQsPQIRAsPsPQPssQPLQIHRQtPEsKNA
Q9NPI6 S523 Sugiyama DCP1A SMIF APSVFQQTVTRSSDLERKAssPsPLtIGtPEsQRKPSIILS
Q9P265 T71 Sugiyama DIP2B KIAA1463 PysPQTQETDSAVQKELRNQtPAPsAAQtsAPsKYHRTRSG
Q9UER7 S668 SIGNOR DAXX BING2 DAP6 YVERQRSVHEKNGKKICtLPsPPsPLAsLAPVADsstRVDs
Q9UGI8 S214 Sugiyama TES LPCEMDAQGPKQMNIPGGDRstPAAVGAMEDKSAEHKRTQY
Q9UMZ2 S1087 Sugiyama SYNRG AP1GBP1 SYNG QGSHKRsLsLGDKEIsRssPsPALEQPFRDRsNtLNEKPAL
Q9UQ35 S1103 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 sPsLQSKSQTsPKGGRsRsssPVtELAsRsPIRQDRGEFsA
Q9UQ35 S2132 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 SRMsCFsRPsMsPtPLDRCRsPGMLEPLGssRtPMsVLQQA
Q9UQ35 S323 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 RGEGDAPFsEPGttstQRPssPEtAtKQPssPYEDKDKDKK
Q9UQ35 T318 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 PAsGRRGEGDAPFsEPGttstQRPssPEtAtKQPssPYEDK
Q9UQ35 T328 Sugiyama SRRM2 KIAA0324 SRL300 SRM300 HSPC075 APFsEPGttstQRPssPEtAtKQPssPYEDKDKDKKEKsAt
Q9Y2B0 S65 Sugiyama CNPY2 MSAP TMEM4 ZSIG9 UNQ1943/PRO4426 VDPKKTIQMGsFRINPDGsQsVVEVPyARsEAHLtELLEEI
Q9Y2U5 S163 Sugiyama MAP3K2 MAPKKK2 MEKK2 VFGAERKKRLsIIGPTSRDRssPPPGYIPDELHQVARNGSF
Q9Y2U5 S164 Sugiyama MAP3K2 MAPKKK2 MEKK2 FGAERKKRLsIIGPTSRDRssPPPGYIPDELHQVARNGSFT