LOK

All methods over the ±5 window, side by side.

PSPA

CDDM

MLP-attr

Normalized values from positional scanning peptide array

PSPA measures pT, so at flanking positions the pS row is a duplicate of the pT row. At position 0 the acceptor identity (pS, pT, pY) is genuine.

Position-wise Probabilities

Log-Odds: Probabilities / STY Background

Position-wise Probabilities (±5 window)

Log-Odds (±5 window)

Motifs stratified by phospho-acceptor (S / T / Y)

K-means sub-motif clusters

Signed attribution PSSM distilled from the CDDM discriminative MLP (in-silico saturation mutagenesis). Letters above 0 are enriched, below are depleted; this is not a probability distribution.

MLP-attribution PSSM (±5, signed)

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substrate_uniprot	site	source	substrate_genes	site_seq
O43615	T106	Sugiyama	TIMM44 MIMT44 TIM44	DEARRLEEsDVLQEARRKyKtIESETVRTSEVLRKKLGELT
O43615	Y104	Sugiyama	TIMM44 MIMT44 TIM44	FRDEARRLEEsDVLQEARRKyKtIESETVRTSEVLRKKLGE
O75534	T761	Sugiyama	CSDE1 D1S155E KIAA0885 NRU UNR	GPKAVAAPRPDRLVNRLKNItLDDAsAPRLMVLRQPRGPDN
O75821	T240	Sugiyama	EIF3G EIF3S4	DGAsRRGESMQPNRRADDNAtIRVtNLSEDTRETDLQELFR
O94804	S438	Sugiyama	STK10 LOK	MDARIQVAQEKQVAEQGGDLsPAANRsQKAsQsRPNssALE
O94804	S444	Sugiyama	STK10 LOK	VAQEKQVAEQGGDLsPAANRsQKAsQsRPNssALEtLGGEK
O94804	S448	Sugiyama	STK10 LOK	KQVAEQGGDLsPAANRsQKAsQsRPNssALEtLGGEKLANG
O94804	S450	Sugiyama	STK10 LOK	VAEQGGDLsPAANRsQKAsQsRPNssALEtLGGEKLANGsL
O94804	S454	Sugiyama	STK10 LOK	GGDLsPAANRsQKAsQsRPNssALEtLGGEKLANGsLEPPA
O94804	S455	iPTMNet\|EPSD\|Sugiyama	STK10 LOK	GDLsPAANRsQKAsQsRPNssALEtLGGEKLANGsLEPPAQ
O94804	S469	Sugiyama	STK10 LOK	QsRPNssALEtLGGEKLANGsLEPPAQAAPGPSKRDsDCss
O94804	S541	Sugiyama	STK10 LOK	KLYKKTLKRTRKFVVDGVEVsITtsKIISEDEKKDEEMRFL
O94804	S545	Sugiyama	STK10 LOK	KTLKRTRKFVVDGVEVsITtsKIISEDEKKDEEMRFLRRQE
O94804	S824	Sugiyama	STK10 LOK	ARLPKIQRSEGKTRMAMYKKsLHINGGGSAAEQREKIKQFS
O94804	S954	Sugiyama	STK10 LOK	EQEMFFKLSEEAECPNPstPsKAAKFFPySSADAs______
O94804	S968	Sugiyama	STK10 LOK	PNPstPsKAAKFFPySSADAs____________________
O94804	T459	Sugiyama	STK10 LOK	PAANRsQKAsQsRPNssALEtLGGEKLANGsLEPPAQAAPG
O94804	T544	Sugiyama	STK10 LOK	KKTLKRTRKFVVDGVEVsITtsKIISEDEKKDEEMRFLRRQ
O94804	T611	Sugiyama	STK10 LOK	LEQMHKRFEQEINAKKKFFDtELENLERQQKQQVEKMEQDH
O94804	T952	Sugiyama	STK10 LOK	KREQEMFFKLSEEAECPNPstPsKAAKFFPySSADAs____
O94804	Y766	Sugiyama	STK10 LOK	MEEHQLQERHQLVKQQLKDQyFLQRHELLRKHEKEREQMQR
P00338	T322	Sugiyama	LDHA PIG19	DLVKVTLtsEEEARLKKsADtLWGIQKELQF__________
P00558	T168	Sugiyama	PGK1 PGKA MIG10 OK/SW-cl.110	EAFRAsLSKLGDVyVNDAFGtAHRAHssMVGVNLPQKAGGF
P04075	S39	Sugiyama	ALDOA ALDA	IAHRIVAPGKGILAADEstGsIAKRLQsIGtENtEENRRFy
P04406	T246	Sugiyama	GAPDH GAPD CDABP0047 OK/SW-cl.12	GKLtGMAFRVPtANVsVVDLtCRLEKPAKyDDIKKVVKQAS
P04792	S82	Sugiyama	HSPB1 HSP27 HSP28	AIEsPAVAAPAYsRALsRQLssGVsEIRHtADRWRVsLDVN
P05455	S325	Sugiyama	SSB	VLEGEVEKEALKKIIEDQQEsLNKWKSKGRRFKGKGKGNKA
P07195	T323	Sugiyama	LDHB	sVINQKLKDDEVAQLKKsADtLWDIQKDLKDL_________
P07205	T168	Sugiyama	PGK2 PGKB	EAFRAsLSKLGDVyVNDAFGtAHRAHSSMVGVNLPHKASGF
P07814	T1506	Sugiyama	EPRS1 EPRS GLNS PARS QARS QPRS PIG32	LCELQPGAKCVCGKNPAKyytLFGRsy______________
P08238	S462	Sugiyama	HSP90AB1 HSP90B HSPC2 HSPC3 HSPCB	HEDstNRRRLsELLRyHtsQsGDEMtsLsEyVsRMKEtQKs
P08238	T637	Sugiyama	HSP90AB1 HSP90B HSPC2 HSPC3 HSPCB	MGyMMAKKHLEINPDHPIVEtLRQKAEADKNDKAVKDLVVL
P09382	T58	Sugiyama	LGALS1	DsNNLCLHFNPRFNAHGDANtIVCNsKDGGAWGTEQREAVF
P0DPH7	T334	Sugiyama	TUBA3C TUBA2	ACCMLYRGDVVPKDVNAAIAtIKtKRTIQFVDWCPTGFKVG
P11142	T163	Sugiyama	HSPA8 HSC70 HSP73 HSPA10	VVtVPAyFNDsQRQAtKDAGtIAGLNVLRIINEPtAAAIAy
P11142	T66	Sugiyama	HSPA8 HSC70 HSP73 HSPA10	DtERLIGDAAKNQVAMNPtNtVFDAKRLIGRRFDDAVVQsD
P14618	T50	Sugiyama	PKM OIP3 PK2 PK3 PKM2	MCRLDIDsPPItARNtGIICtIGPAsRsVEtLKEMIKsGMN
P15311	T567	EPSD\|PSP	EZR VIL2	RTHNDIIHNENMRQGRDKYKtLRQIRQGNTKQRIDEFEAL_
P26038	T558	GPS6\|SIGNOR\|ELM\|EPSD\|PSP	MSN	KTANDMIHAENMRLGRDKYKtLRQIRQGNTKQRIDEFEsM_
P26373	S181	Sugiyama	RPL13 BBC1 OK/SW-cl.46	YKKEKARVItEEEKNFKAFAsLRMARANARLFGIRAKRAKE
P26641	T287	Sugiyama	EEF1G EF1G PRO1608	EQALAAEPKAKDPFAHLPKstFVLDEFKRKysNEDtLsVAL
P31948	T243	Sugiyama	STIP1	KQALKEKELGNDAYKKKDFDtALKHYDKAKELDPtNMtyIT
P35241	T564	EPSD\|PSP	RDX	KTQNDVLHAENVKAGRDKYKtLRQIRQGNtKQRIDEFEAM_
P35269	T41	Sugiyama	GTF2F1 RAP74	NTTKKyNIMAFNAADKVNFAtWNQARLERDLSNKKIyQEEE
P37802	T180	Sugiyama	TAGLN2 KIAA0120 CDABP0035	RNFsDNQLQEGKNVIGLQMGtNRGAsQAGMtGyGMPRQIL_
P38646	T116	Sugiyama	HSPA9 GRP75 HSPA9B mt-HSP70	DGERLVGMPAKRQAVtNPNNtFyAtKRLIGRRYDDPEVQKD
P38646	T120	Sugiyama	HSPA9 GRP75 HSPA9B mt-HSP70	LVGMPAKRQAVtNPNNtFyAtKRLIGRRYDDPEVQKDIKNV
P38646	Y118	Sugiyama	HSPA9 GRP75 HSPA9B mt-HSP70	ERLVGMPAKRQAVtNPNNtFyAtKRLIGRRYDDPEVQKDIK
P39019	Y54	Sugiyama	RPS19	VDtVKLAKHKELAPyDENWFytRAAstARHLYLRGGAGVGs
P47756	T202	Sugiyama	CAPZB	SGsGtMNLGGsLtRQMEKDEtVsDCsPHIANIGRLVEDMEN
P53350	T210	GPS6\|ELM\|EPSD\|PSP	PLK1 PLK	VKIGDFGLATKVEYDGERKKtLCGtPNyIAPEVLSKKGHSF
P55209	T269	Sugiyama	NAP1L1 NRP	GPEIMGCTGCQIDWKKGKNVtLKtIKKKQKHKGRGTVRTVT
P55209	T272	Sugiyama	NAP1L1 NRP	IMGCTGCQIDWKKGKNVtLKtIKKKQKHKGRGTVRTVTKTV
P60709	S52	Sugiyama	ACTB	PsIVGRPRHQGVMVGMGQKDsyVGDEAQsKRGILTLKYPIE
P61254	T8	Sugiyama	RPL26	_____________MKFNPFVtsDRsKNRKRHFNAPsHIRRK
P62280	T15	Sugiyama	RPS11	______MADIQTERAyQKQPtIFQNKKRVLLGEtGKEKLPR
P62701	T247	Sugiyama	RPS4X CCG2 RPS4 SCAR	VIGKGNKPWIsLPRGKGIRLtIAEERDKRLAAKQSSG____
P62736	S54	Sugiyama	ACTA2 ACTSA ACTVS GIG46	PsIVGRPRHQGVMVGMGQKDsyVGDEAQsKRGILTLKyPIE
P62899	T26	Sugiyama	RPL31	KGGEKKKGRsAINEVVtREytINIHKRIHGVGFKKRAPRAL
P62917	T135	Sugiyama	RPL8	CLEEKPGDRGKLARAsGNyAtVIsHNPEtKKtRVKLPSGSK
P62917	Y133	Sugiyama	RPL8	VCCLEEKPGDRGKLARAsGNyAtVIsHNPEtKKtRVKLPSG
P62942	T7	Sugiyama	FKBP1A FKBP1 FKBP12	______________MGVQVEtIsPGDGRtFPKRGQtCVVHy
P63261	S52	Sugiyama	ACTG1 ACTG	PsIVGRPRHQGVMVGMGQKDsyVGDEAQsKRGILTLKYPIE
P63267	S53	Sugiyama	ACTG2 ACTA3 ACTL3 ACTSG	PsIVGRPRHQGVMVGMGQKDsyVGDEAQsKRGILTLKyPIE
P68032	S54	Sugiyama	ACTC1 ACTC	PsIVGRPRHQGVMVGMGQKDsyVGDEAQsKRGILTLKyPIE
P68133	S54	Sugiyama	ACTA1 ACTA	PsIVGRPRHQGVMVGMGQKDsyVGDEAQsKRGILTLKyPIE
P68363	T334	Sugiyama	TUBA1B	ACCLLyRGDVVPKDVNAAIAtIKtKRsIQFVDWCPTGFKVG
Q14697	T40	Sugiyama	GANAB G2AN KIAA0088	LAFLGVCLGITLAVDRSNFKtCEEssFCKRQRsIRPGLsPy
Q6PEY2	T334	Sugiyama	TUBA3E	ACCMLYRGDVVPKDVNAAIAtIKtKRTIQFVDWCPTGFKVG
Q71U36	T334	Sugiyama	TUBA1A TUBA3	ACCLLyRGDVVPKDVNAAIAtIKtKRTIQFVDWCPTGFKVG
Q8TD47	T247	Sugiyama	RPS4Y2 RPS4Y2P	VIGNGNKPWISLPRGKGIRLtIAEERDKRLAAKQSSG____
Q99426	T120	Sugiyama	TBCB CG22 CKAP1	DVSRVEKytIsQEAyDQRQDtVRSFLKRsKLGRyNEEERAQ
Q9BQE3	T334	Sugiyama	TUBA1C TUBA6	ACCLLyRGDVVPKDVNAAIAtIKtKRTIQFVDWCPTGFKVG
Q9H173	T125	Sugiyama	SIL1 UNQ545/PRO836	QYEDKFRNNLKGKRLDINtNtytsQDLKSALAKFKEGAEME
Q9ULW6	T342	Sugiyama	NAP1L2 BPX	GTAIEYSTGCEIDWNEGKNVtLKtIKKKQKHRIWGTIRTVT
Q9ULW6	T345	Sugiyama	NAP1L2 BPX	IEYSTGCEIDWNEGKNVtLKtIKKKQKHRIWGTIRTVTEDF
Q9UNX3	T8	Sugiyama	RPL26L1 RPL26P1	_____________MKFNPFVtsDRsKNRKRHFNAPSHVRRK
Q9UPQ0	S377	Sugiyama	LIMCH1 KIAA1102	LREEDDKWQDDLARWKSRRRsVsQDLIKKEEERKKMEKLLA
Q9Y597	T673	Sugiyama	KCTD3	sPLLARARRtEsFHSYRDFQtINLNRNVERAVPENGNLGPI